Ahrari, A., Sharifi, A. and Haghighi, A. T. (2024). Anthropogenic vs. climatic drivers: Dissecting Lake desiccation on the Iranian plateau. Journal of Environmental Management, 368: 122103. https://doi.org/10.1016/j.jenvman.2024.122103
Çiçek, E., Esmaeili, H. R., Sayyadzadeh, G., Saad, A., Sungur, S., Jawad, L., Eagderi, S., Çapar, O. B., Coad, B. W. and Fricke, R. (2024). Freshwater lampreys and fishes in the Middle East. Taxa (Journal of Taxonomy and Systematics), 4: 1–429.
Çiçek, E., Jawad, L., Sungur, S., Eagderi, S., Esmaeili, H. R., Mouludi-Saleh, A. and Fricke, R. (2023). Freshwater fishes of Iraq: a revised and updated annotated checklist - 2023. Zootaxa, 5357 (1): 1–49. https://doi.org/10.11646/zootaxa.5357.1.1
Coad, B. W. (1995). Freshwater fishes of Iran. Acta Scientiarum Naturalium Academiae Scientiarum Bohemicae, Brno, 29 (1): 1–64.
Dudgeon, D., Arthington, A. H., Gessner, M. O., Kawabata, Z. I., Knowler, D. J., Lévêque, C., Naiman, R. J., Prieur-Richard, A. H., Soto, D., Stiassny, M. L. J. and Sullivan, C. A. (2006). Freshwater biodiversity: importance, threats, status and conservation challenges. Biological Reviews, 81 (2): 163–182. https://doi.org/10.1017/S1464793105006950
Eagderi, S., Mouludi-Saleh, A., Esmaeili, H. R., Sayyadzadeh, G. and Nasri, M. (2022). Freshwater lamprey and fishes of Iran; a revised and updated annotated checklist - 2022. Turkish Journal of Zoology, 46 (6): 500–522. https://doi.org/10.55730/1300-0179.3104
Esmaeili, H. R., Coad, B. W., Gholamifard, A., Nazari, N. and Teimory, A. (2010). Annotated checklist of the freshwater fishes of Iran. Zoosystematica Rossica, 19: 361–386. https://doi.org/10.31610/zsr/2010.19.2.361
Esmaeili, H. R., Mehraban, H., Abbasi, K., Keivany, Y. and Coad, B. W. (2017). Review and updated checklist of freshwater fishes of Iran: taxonomy, distribution and conservation status. Iranian Journal of Ichthyology, 4 (Suppl. 1): 1–114. https://doi.org/10.22034/iji.v4iSuppl.1.220
Esmaeili, H. R., Sayyadzadeh, G., Eagderi, S. and Abbasi, K. (2018). Checklist of freshwater fishes of Iran. FishTaxa, 3 (3): 1–95.
Esmaeili, H. R., Sayyadzadeh, G., Masoumi, A. H., Hashemi, S. H. and Echreshavi, S. (2025). Tariqilabeo iranicus, a new species of labeonine fish from the Makran basin in Iran (Teleostei: Cyprinidae). Zootaxa, 5620 (3): 437–450. https://doi.org/10.11646/zootaxa.5620.3.3
Esmaeili, H. R., Sayyadzadeh, G., Zarei, F., Eagderi, S. and Mousavi-Sabet, H. (2022). Mystus cyrusi, a new species of bagrid catfish (Teleostei: Bagridae) from Middle East. Zootaxa, 5099 (3): 325–343. https://doi.org/10.11646/zootaxa.5099.3.2
Freyhof, J. and Yoğurtçuoğlu, B. (2020). A proposal for a new generic structure of the killifish family Aphaniidae, with the description of Aphaniops teimorii (Teleostei: Cyprinodontiformes). Zootaxa, 4810 (3): 421–451. https://doi.org/10.11646/zootaxa.4810.3.2
Freyhof, J., Esmaeili, H. R., Sayyadzadeh, G. and Geiger, M. (2014). Review of the crested loaches of the genus Paracobitis from Iran and Iraq with the description of four new species (Teleostei: Nemacheilidae). Ichthyological Exploration of Freshwaters, 25 (1): 11–38.
Freyhof, J., Kaya, C. and Ali, A. (2021). A critical checklist of the inland fishes native to the Euphrates and Tigris drainages. In: Jawad, L. A. (Ed.), Tigris and Euphrates Rivers: Their Environment from Headwaters to Mouth. Aquatic Ecology Series, Vol. 11. Springer, Cham. pp. 815–854. https://doi.org/10.1007/978-3-030-57570-0_35
Freyhof, J., Segherloo, I. H., Vatandoust, S., Abdollahi-Mousavi, S. E., Normandeau, E., Geiger, M. F. and Yoğurtçuoğlu, B. (2025). Resolving a 182-year-old taxonomic puzzle: Luciobarbus in the Persian Gulf basin (Teleostei: Cyprinidae). Zootaxa, 5620 (1): 29–71. https://doi.org/10.11646/zootaxa.5620.1.3
Fricke, R., Eschmeyer, W. N. and van der Laan, R. (editors) (2026). Eschmeyer’s Catalog of Fishes: Genera, Species, References. Electronic version (Accessed 10 March 2026). http://researcharchive.calacademy.org/research/ichthyology/catalog/fishcatmain.asp
Heckel, J. J. (1846–1849b). Anhang. Die Fische Persiens gesammelt von Theodor Kotschy, In: Russegger, J. (Ed.), Reisen in Europa, Asien und Afrika, mit besonderer Rücksicht auf die naturwissenschaftlichen Verhältnisse der betreffenden Länder, unternommen in den Jahren 1835 bis 1841 von Joseph Russegger. Schweitzerbart'sche Verlagsbuchhandlung, Stuttgart, 2 (3): 255–272.
IUCN (2026). The IUCN Red List of Threatened Species. Version 2025-2. https://www.iucnredlist.org (Accessed 112 March 2026).
Jokar, M., Kamangar, B. B., Ghaderi, E. and Freyhof, J. (2023). Glyptothorax sardashtensis, a new species of torrent catfish from the upper Lesser Zab drainage in Iran (Teleostei: Sisoridae). Zootaxa, 5254 (4): 476–492. https://doi.org/10.11646/zootaxa.5254.4.2
Jouladeh-Roudbar, A., Ghanavi, H. R. and Doadrio, I. (2020). Ichthyofauna from Iranian freshwater: annotated checklist, diagnosis, taxonomy, distribution and conservation assessment. Zoological Studies, 59: 21. https://doi.org/10.6620/ZS.2020.59-21
Jouladeh-Roudbar, A., Ghanavi, H. R. and Freyhof, J. (2023a). Glyptothorax vatandousti, a new species of torrent catfish from the upper Karkheh drainage in Iran (Teleostei: Sisoridae). Zootaxa, 5315 (1): 37–58. https://doi.org/10.11646/zootaxa.5315.1.2
Jouladeh-Roudbar, A., Kaya, C., Vatandoust, S. and Ghanavi, H. R. (2024). New insights into the phylogeny of Carasobarbus Karaman, 1971 (Actinopterygii, Cyprinidae) with the description of three new species. Scientific Reports, 14: 21801. https://doi.org/10.1038/s41598-024-71463-7
Jouladeh-Roudbar, A., Vatandoust, S., Doadrio, I. and Ghanavi, H. R. (2023b). Integrative taxonomy of Turcinoemacheilus Bănărescu & Nalbant, 1964 in West Asia with the description of three new species (Teleostei: Nemacheilidae). Diversity, 15 (12): 1222. https://doi.org/10.3390/d15121222
Khaefi, R., Esmaeili, H. R., Sayyadzadeh, G., Geiger, M. F. and Freyhof, J. (2016). Squalius namak, a new chub from Lake Namak basin in Iran (Teleostei: Cyprinidae). Zootaxa, 4169 (1): 145–159. https://doi.org/10.11646/zootaxa.4169.1.7
Mayden, R. L. (1997). A hierarchy of species concepts: the denouement in the saga of the species problem. In: Claridge, M. F., Dawah, H. A. and Wilson, M.R. (eds.), Species: The Units of Biodiversity. Chapman and Hall, London. pp. 381–424.
Myers, N., Mittermeier, R. A., Mittermeier, C. G., da Fonseca, G. A. B. and Kent, J. (2000). Biodiversity hotspots for conservation priorities. Nature, 403: 853–858. https://doi.org/10.1038/35002501
Padial, J. M., Miralles, A., De la Riva, I. and Vences, M. (2010). The integrative future of taxonomy. Frontiers in Zoology, 7: 16. https://doi.org/10.1186/1742-9994-7-16
Sayyadzadeh, G. and Esmaeili, H. R. (2024). Freshwater lamprey and fishes of Iran: reappraisal and updated checklist with a note on Eagderi et al. (2022). Zootaxa, 5402 (1): 1–99. https://doi.org/10.11646/zootaxa.5402.1.1
Sayyadzadeh, G. and Esmaeili, H.R. (2020). Oxynoemacheilus marunensis, a new loach species from the Persian Gulf basin with remarks on O. frenatus (Teleostei: Nemacheilidae). Zootaxa, 4885 (2): 189–206. https://doi.org/10.11646/zootaxa.4885.2.2
Sayyadzadeh, G., Eagderi, S. and Esmaeili, H. R. (2016). A new loach of the genus Oxynoemacheilus from the Tigris River drainage and its phylogenetic relationships among the nemacheilid fishes (Teleostei: Nemacheilidae) in the Middle East based on mtDNA COI sequences. Iranian Journal of Ichthyology, 3 (4): 236–250.
Sayyadzadeh, G., Teimori, A. and Esmaeili, H. R. (2019). Paraschistura kermanensis, a new stone loach species from southeastern Iran (Teleostei: Nemacheilidae). Zootaxa, 4638 (4): 571–583. https://doi.org/10.11646/zootaxa.4638.4.7
Sayyadzadeh, G., Zarei, F. and Esmaeili, H. R. (2022). Glyptothorax (Teleostei: Sisoridae) from the Middle East: an integrated molecular and morphological insight into its taxonomic diversity. Diversity, 14: 884. https://doi.org/10.3390/d14100884
Schwarzer, J., Shabani, N., Esmaeili, H. R., Mwaiko, S. and Seehausen, O. (2016). Allopatric speciation in the desert - diversification of cichlids at their geographical and ecological range limit in Iran. Hydrobiologia, 791: 193–207. https://doi.org/10.1007/s10750-016-2976-3
Zamani-Faradonbe, M., Zhang, E. and Keivany, Y. (2021). Garra hormuzensis, a new species from the upper Kol River drainage in the Persian Gulf basin (Teleostei: Cyprinidae). Zootaxa, 5052 (3): 380–394. https://doi.org/10.11646/zootaxa.5052.3.4
Zarei, F., Esmaeili, H. R., Kovačić, M., Schliewen, U. K. and Abbasi, K. (2022). Ponticola hircaniaensis sp. nov., a new and critically endangered gobiid species (Teleostei: Gobiidae) from the southern Caspian Sea basin. Zootaxa, 5154 (4): 401–430. https://doi.org/10.11646/zootaxa.5154.4.1
