Ahmad, S., Hameed, S., Ali, H., Khan, T. U., Mehmood, T. and Nawaz, M. A. (2016). Carnivores’ diversity and conflicts with humans in Musk Deer National Park, Azad Jammu and Kashmir, Pakistan. European Journal of Wildlife Research, 62 (5): 565–576.
Andheria, A. P., Karanth, K. U. and Kumar, N. S. (2007). Diet and prey profiles of three sympatric large carnivores in Bandipur Tiger Reserve, India. Journal of Zoology, 273 (2): 169–175.
Araujo, M. B. and Guisan, A. (2006) Five (or so) challenges for species distribution modelling. Journal of Biogeography, 33 (10): 1677–1688.
Banerjee, A. and Bandopadhyay, R. (2016). Biodiversity hotspot of Bhutan and its sustainability. Current Science, 110 (4): 521.
Bhatt, U., Adhikari, B. and Lyngdoh, S. (2022). Monitoring diversity and abundance of mammals with camera-traps: a case study of Manas National Park, Assam, India. Check List, 18 (5): 1023–1043.
BTF. (2023). Bhutan Trust Fund. 2023. https://bhutantrustfund.bt/public_page/post_details/18 (Accessed 10 January, 2024).
BWS. (2020). Conservation Management Plan of Bumdeling Wildlife Sanctuary (July 2020-June 2030). Department of Forests and Park Services, Ministry of Energy and Natural Resources, Royal Government of Bhutan. 12 pp.
Carbone, C., Christie, S., Conforti, K., Coulson, T., Franklin, N., Ginsberg, J. R., Griffiths, M., Holden, J., Kawanishi, K., Kinnaird, M. and Laidlaw, R. (2001). The use of photographic rates to estimate densities of tigers and other cryptic mammals. Animal Conservation Forum, 4 (1): 75–79.
CITES. (2025). Convention on International Trade in Endangered Species of Wild Fauna and Flora (CITES). (2025). Appendices I, II and III. Retrieved from https://cites.org/eng/app/appendices.php (Accessed 9 January 2025).
Dhendup, T. and Letro. (2016). Livestock depredation in Jigme Singye Wangchuk National Park. NeBio, an International Journal of Environment and Biodiversity, 7 (3): 78–81.
Dhendup, T. and Tamang, K. (2023). Rich felid diversity in forests around Thimphu, Bhutan’s capital city. Cat News, 76: 43–45.
Dhendup, T., Thinley, K. and Tenzin, U. (2019). Mammal diversity in a montane forest in central Bhutan. Journal of Threatened Taxa, 11 (13): 14757–14763.
DoFPS. (2016). National Snow Leopard Survey of Bhutan 2014–2016 (Phase II): Camera Trap Survey for Population Estimation. Department of Forests and Park Services, Ministry of Agriculture and Forests, Thimphu, Bhutan. 9 pp.
Estes, J. A., Terborgh, J., Brashares, J. S., Power, M. E., Berger, J., Bond, W. J., Carpenter, S. R., Essington, T. E., Holt, R. D., Jackson, J. B. C., Marquis, R. J., Oksanen, L., Oksanen, T., Paine, R. T., Pikitch, E. K., Ripple, W. J., Sandin, S. A., Scheffer, M., Schoener, T. W., Shurin, J. B., Sinclair, A. R. E., Soulé, M. E., Virtanen, R. and Wardle, D. A. (2011). Trophic downgrading of planet Earth. Science, 333 (6040): 301–306.
FMID. (2023). Forest Monitoring and Information Division. 2023. National Forest Inventory Volume I: State of Forest Report. Department of Forest and Park Services. 9 pp.
FNCA. (2023). Forest and Nature Conservation Act of Bhutan. 2023. 67 pp.
IUCN. (2025). International Union for Conservation of Nature (IUCN) Red List of Threatened Species. Version 2025. https://www.iucnredlist.org (Accessed 9 January 2025).
JWS. (2023). Jomotsangkha Wildlife Sanctuary, 2023. Conservation Management Plan, Department of Forests and Park Services, Ministry of Energy and Natural Resources, Royal Government of Bhutan. 26 pp.
Karanth, K. U., Nichols, J. D., Kumar, N. S., Link, W. A. and Hines, J. E. (2004). Tigers and their prey: Predicting carnivore densities from prey abundance. Proceedings of the National Academy of Sciences of the United States of America, 101 (14): 4854–4858.
Layrab, S. D. L. (2023). Livestock depredation and its impact on farmers in Trongsa district, Bhutan. Indonesian Journal of Social and Environmental Issues, 4 (2): 192–201.
Letro, L., Duba, D., Tandin, T. and Wangdi, S. (2021). Rare capture of a snow leopard in Jigme Singye Wangchuck National Park, Bhutan. Cat News, 73: 29–32.
Linnell, J. D., Andersen, R., Kvam, T., Andren, H., Liberg, O., Odden, J. and Moa, P. (2001). Home range size and choice of management strategy for lynx in Scandinavia. Environmental Management, 27 (6): 869–879.
Macdonald, D. W. and Sillero-Zubiri, C. (2002). Large carnivores and conflict: Lion conservation in context, In: Loveridge, A. J., Lynam, T. and Macdonald, D. W. (Eds.), Lion conservation research. Workshop 2: modelling conflict. Wildlife Conservation Research Unit, Oxford University. pp. 1–8.
Naseer, A., Bilal, M., Naseer, U., Mustafa, N. and Rakha, B. A. (2020). Population density, habitat characteristics and preferences of Red Fox (Vulpes vulpes) in Chakwal, Pakistan. Journal of Bioresource Management, 7 (4): 74–83.
NCD. (2008). Nature Conservation Division. (2008). Bhutan National Human-wildlife Conflicts Management Strategy, Department of Forest and Park Services, Ministry of Agriculture. 4 pp.
O’Brien, T. G., Kinnaird, M. F. and Wibisono, H. T. (2003). Crouching tigers, hidden prey: Sumatran tiger and prey populations in a tropical forest landscape. Animal Conservation, 6 (2): 131–139.
https://doi.org/10.1017/s1367943003003172
Pringle, R. M., Naiman, R. J., Breithaupt, J. L., Eisenhauer, N., Jackson, B. J. and Manning, D. W. P. (2017). Upward cascades in rivers: Do salmon subsidize terrestrial wildlife in the Pacific Northwest? Bioscience, 67 (7): 676–686.
Rabinowitz, A. R. and Walker, R. S. (1991). The carnivore community in a dry tropical forest mosaic in Huai Kha Khaeng Wildlife Sanctuary, Thailand. Journal of Tropical Ecology, 7 (1): 37–47.
Ramírez-Álvarez, D., Napolitano, C., Arriagada, G., Salgado, I., Cox, S. and Céspedes-Parada, B. (2023). Native carnivore diversity and relative abundance in landscapes of the coast range in Central Chile: Insights for Conservation Decision-Making. Conservation, 3 (3): 379–393.
Ripple, W. J., Estes, J. A., Beschta, R. L., Wilmers, C. C., Ritchie, E. G., Hebblewhite, M., Berger, J., Elmhagen, B., Letnic, M., Nelson, M. P., Schmitz, O. J., Smith, D. W., Wallach, A. D. and Wirsing, A. J. (2014). Status and ecological effects of the world's largest carnivores. Science, 343 (6167): 151–163.
Shao, X., Lu, Q., Xiong, M., Bu, H., Shi, X., Wang, D., Zhao, J., Li, S. and Yao, M. (2021). Prey partitioning and livestock consumption in the world’s richest large carnivore assemblage. CB/Current Biology, 31 (22): 4887–4897.e5.
Shrestha, L., Sarkar, M. S., Shrestha, K., Aung, P. S., Wen, X., Yongping, Y., Zhipang, H., Xuefei, Y., Shaoliang, Y. and Chettri, N. (2022). Mammalian research, diversity and conservation in the Far Eastern Himalaya Landscape: A review. Global Ecology and Conservation, 34: e02003.
Shrotriya, S., Reshamwala, H. S., Lyngdoh, S., Jhala, Y. V. and Habib, B. (2022). Feeding patterns of three widespread Carnivores—The wolf, snow leopard, and Red Fox—in the Trans-Himalayan landscape of India. Frontiers in Ecology and Evolution, 10: 815996.
Soberón J. (2007). Grinnellian and Eltonian niches and geographic distributions of species. Ecology Letters, 10 (12): 1115–1123.
SWS. (2019). Conservation Management Plan of Sakteng Wildlife Sanctuary (Amendment, 2019). Department of Forest and Park Services, Royal Government of Bhutan. 7 pp.
Vernes, K., Rajaratnam, R. and Dorji, S. (2022). Patterns of species co‑ occurrence in a diverse Eastern Himalayan montane carnivore community. Mammal Research, 67 (2): 139–149.
Wangchuk, T., Thinley, P., Tshering, K., Tshering, C., Yonton, D. and Pema, B. (2004). Mammals of Bhutan. Department of Forest and Park Services, Thimphu, Bhutan. 180 pp.
Wangdi, T., Tobgay, S., Dorjee, K., Dorji, K. and Wangyel, S. (2019). The distribution, status and conservation of the Himalayan Musk Deer Moschus chrysogaster in Sakteng Wildlife Sanctuary. Global Ecology and Conservation, 17: e00466.
WWF. (2023). Biological Corridor - 9: A green ribbon between Sakteng Wildlife Sanctuary and Bumdeling Wildlife Sanctuary. https://www.wwfbhutan.org.bt/?381915/Biological-corridor-9. (Accessed 1 October 2024).
ZFD. (2023). Zhemgang Forest Division. 2023. Conservation Management Plan of Biological Corridor 4 (January 2023 to December 2032). Department of Forests and Park Services, Ministry of Agriculture and Forests, Royal Government of Bhutan, Zhemgang. 2 pp.